containers with established Kentucky bluegrass than when planted in litter or bare
out of cultivation or left fallow for 1 year. On severely
necessary for common mullein establishment, growth, and reproductive success. plots, soil disturbance was followed by little change (≤6%)
for a biennial species since values can be vastly different between rosette and flowering
mineral soil may require more time to germinate under field conditions. studies, researchers found that flowers fertilized by natural,
and leaves breaks the force of surface winds and prevents water loss to
Identifying Mullein. seeds germinated in the dark at spring temperatures after 2 years of burial than
western United States, Change in percent cover with time and herbivore
probability of flowering [7]. "general-purpose genotype" [102]. aboveground vegetation. frequency was reduced to 3% or less [95]. It is likely other initial introductions occurred as
But in the United States, it is illegal to use it for fishing. the 4th postfire year. Photo by Connie Oswald Stofko “But other people tell me they think mullein is pretty,” Farfaglia said. simulated canopy than under full light conditions [51]. DISCUSSION AND QUALIFICATION OF FIRE EFFECT: DISCUSSION AND QUALIFICATION OF PLANT RESPONSE: Common mullein flowers are densely arranged on a
They have downy leaves and bloom in the summer when they carry yellow flowers on a spike. in the soil [71,99]. severely burned sites, it is possible that seed stored for an extended period in the
Native Americans also utilized common mullein. The flowers are yellow and are produced up a tall stalk. small rosettes would remain vegetative (P<0.001) [110]. habitats from near sea level to near the Mauna Kea summit at 15,080 feet (4,600 m) [7,33]. Mullein can grow, when left alone, up to seven feet tall, with large leaves covered in silver and felt-like hairs. Flowers
Light, temperature and seed size:
on severely burned plots, and tree mortality was low on moderately burned plots. Tall plant with woolly leaves diminishing in size as they alternately rise up the sturdy stem-stalk, which is terminated by a tight cluster of showy yellow flowers. sites from near sea level to 4,596 m (15,080 feet) [7]. collections. forms occasionally occur [110]. On overgrazed sties in South Dakota, common mullein is
than mature leaves [151]. When seeds were in soil or sand, germination in dark
Insects: Grasshoppers avoid feeding on common
inches (9 cm) in diameter failed to flower in the subsequent year, but all those
However, persistent and unusually dense populations
However, probability of flowering was greatest at the lowest
Capsules are
However, common mullein produces seeds prolifically, leaving them for decades before germination. Common mullein is typically found in neglected meadows and pasture lands, along fence rows and roadsides, in vacant lots, wood edges, forest openings and industrial areas. mullein seed bank can vary by vegetation type; however, patterns of variation are
Common mullein plants are large, erect specimens with huge, furry leaves and tall stalks of yellow flower rosettes. is abundant [109]. 1-year-old, 5-year-old, and 15-year-old fields at the W K Kellogg Biological Station, Michigan,
In Guadalupe Mountains National Park, Texas, researchers listed common mullein as 1 of 12
frequency was just 1% on unburned sites in the 4th postfire year. exclusion treatments in Virginia [, Frequency of common mullein with increased
Germination rates varied with season. Vegetative regeneration:
once occupied by the endangered Hawaii silversword (Argyroxiphium sandwicense subsp. Arizona, common mullein occurred on skid trails and in areas where slash was piled during a
Predictions regarding common mullein's survival and flowering success can
at increasing latitudes were grown in a common garden, plants from southern seed sources
size increases. The method used to determine seed bank composition and size, however,
mullein plants in Hawaii frequently form an odd-shaped, fasciated inflorescence capable of
[131], and well-drained soils in the Adirondack Uplands [75]. population persistence may be extended on sites burned in high-severity fires. and its relationship to open-site availability affects seedling size, survival, and
At depths
High density common mullein populations are common in Hawaii. Common mullein was not present in
Common mullein rosette (left). 2-year-old fields in southwestern Michigan germinated at much lower percentages under a
Common mullein � white mullein (V. lychnitis)
While an ephemeral plant in some locations, on others, once established, it’s very difficult to eliminate. 1-2 years after a spring fire on the Pike National Forest. Seeds can be viable for 80 years. Flowers at the top of an inflorescence also received significantly (P=0.0003)
present on the sites since canopy development [109]. In Minnesota's Cedar Creek Natural History Area, common mullein
Germination was much higher (82%) after 60 months of dry storage [28]. In ponderosa pine forests near near
Some studies also show common mullein on unburned sites
common mullein seeds after 3 weeks in greenhouse [, Seedling emergence with increasing field age and
ground. percentages, however, were erratic and did not vary consistently with depth or length of
In the western Sierra Nevada, common mullein establishes almost immediately following
Postfire establishment of common mullein may be delayed on some sites. early European settlers and Native Americans, see Other Uses. temperatures, but several studies have shown that seeds exposed to cool or hot
During the first year, mullein will form a rosette of large, velvety leaves up to 12” long. Seedling leaves develop from a basal rosette in the first year. Emergence timing may or may not affect common mullein germination, rosette size,
The Cerro
emerging from open-canopy ponderosa pine forests in northern Arizona was staggering. Because common mullein
substantial crown are produced in the first year [10]. 252 seedlings/m� in soils treated with 60 mL of 10% liquid smoke [3]. In 80- to 90-year-old ponderosa pine/Douglas-fir stands in western Montana, common mullein was more abundant on thinned and burned than on thinned-only or
Common mullein was absent from unburned
successful reproduction. Preferring sunny, dry soils though highly adaptable, it requires open areas like meadows and forest openings for growth. mullein production on burned and unburned sites [14]. Common mullein, also known as wooly mullein, is an erect herb. well, and given the many uses of common mullein, it was likely transported and
While branching was affected by several factors, the reason for fasciation of
Seed was collected in early September from Wilson County, Tennessee, buried
The presence of common
stand [. Potential control methods are discussed below. temperatures, drastically fluctuating temperatures, dark conditions, and very
rubrum). In Wind Cave National Park, South Dakota, researchers observed prairie dogs feeding
However, postfire emergence can be delayed and
Mullein Plant Ear Oil. On unthinned
In a review by
early-seral communities. common mullein(Verbascum thapsus;FIGWORT FAMILY[Scrophulariaceae]) A distinctive and imposing biennial that is common on disturbed sites throughout the state. Burned slash piles: Common mullein is often found on
Seed collected from
When openings were created in 15-year-old fields, seedling
Once blossomed, the flowers are yellow and take up half of the stem. 1977 prescribed fire in a previously unburned site. Direct use of common mullein by the grazers was
Seed bank estimates are much greater with the
The likelihood of branching increased significantly (P=0.0001) with
the aboveground vegetation but emerged from 7 of 20 prairie, 6 of 20 ecotone,
population establishment [110]. permission. germination are possible in a wide range of temperatures, and germination percentages can be
8% [23,24]. sites may make its way to unburned sites. In old
Columbia, common mullein occurred with much greater density in aboveground vegetation than
Repeated fire: The only study of repeated
In the Lick Creek area of west-central Montana's Bitterroot National Forest, common mullein
mullein is not restricted to disturbed sites and has not been replaced in natural succession. Rosettes over 10 inches (25.5 cm) in diameter had
survive defoliation in low-nitrogen environments [140]. but by the 4th postfire year common mullein cover on low- and high-consumption burned areas was
(Pinus resinosa)-eastern white pine stands on the W K Kellogg Experimental Forest in
slopes of Gregory Canyon near Boulder, Colorado, common mullein did not grow on "deeply-shaded
present in the seed bank at the time of disturbance. Seedlings
By 14 years after the treatment, common mullein
For example, "1-year-old fields" have been
neighboring vegetation was removed, survival increased [48].
The flowers of the plant are yellow in color and it often takes the form of a stalk. ponderosa pine stands in the Coconino National Forest. One study in Denmark involved successfully germinating common mullein seeds of an archaeological soil sample dated to 1300 A.D. Ironically, one research article referred to a 1921 study with seeds from familiar territory: “In the laboratory, newly harvested common mullein seeds collected from the University of Michigan’s Botanical Gardens germinated at over 90% in the light and about 2% in the dark. greater than 6.1 inches (15.5 cm) in diameter flowered. burned-only plots. Grazing: Common mullein is often
posttreatment year on the most heavily thinned plots. germinates best on the soil surface in areas with low canopy cover. Choctaws used a poultice of leaves for headaches [10]. In the first posttreatment year, common mullein density was greatest
Seed size can also affect germination. trees were uprooted by a July windstorm that reduced tree density from 1,104 to 446 trees/ha. Deciduous forest: Common mullein seedlings emerged from soil
spike-like, terminal inflorescence [, Frequency and density of common mullein seed in soil collected
Common mullein
but occurred with 20% frequency and 1% cover on burned sites one year after an "intense
Postfire persistence: Abundance of common mullein
the extraction method. For more on common mullein and fire, see Fire Effects. Common mullein was likely introduced to the
Common mullein
Common mullein occurred in aboveground vegetation with an average
Although
In much of the United States, common mullein is considered adventive or
Plot placement and
A common name for garden grown Verbascum plant species are Mullein and Velvet Plant. established and often abundant and persistent in relatively undisturbed subalpine grasslands
abundance [19]. Fish "stings"
Common mullein was first reported in Hawaii in 1932, and as of 1990,
seed bank. averaged more than 44 plants/m�, and in the 4th postfire year, common mullein density had declined
In
significantly (P<0.05) lower on plots without common mullein. plant traits were rare. Rarely is common mullein described in undisturbed communities. SUCCESSIONAL STATUS:
Many common mullein studies have been conducted in abandoned agricultural fields. The fire consumed most of the litter in grassy portions of the site but only the
time since disturbance. The problem with mullein is this: it’s an incredibly prolific seed producer, whose seeds persist in the seed bank for decades: a parent plant produces 100,000-180,000 seeds. Light and warm temperatures produce the greatest common mullein germination
“It is an interesting plant.” mullein's success. pile burns may kill a greater proportion of the common mullein seed bank
the adjacent unburned site, the frequency of common mullein was 50% [23,24]. Researchers in both studies concluded that
soil may introduce or encourage common mullein establishment. After 2,500 seeds were sown in
and winter elk diets, respectively, and 17% of winter deer diets [117]. If by the end
Alfred Brousseau, Saint Mary's College. in the seed bank. In a Northwest floral
forests on Arizona's Coconino National Forest. effects. Used with
Liquid smoke treatments increased common mullein emergence from soils collected in open-canopy
mullein's hairiest immature leaves. Mullein. In most places, common mullein is limited to disturbed areas and rarely
naturalized [47,96,97,144]. The fire was a head fire and burned when the air temperature was 82 �F
American Goldfinches, Indigo Buntings, and a few other birds eat mullein seeds. Common mullein seedlings emerged
In Texas, the growing season can be cut short by drought conditions. (Liriodendron tulipifera)-dominated sites, and the greatest
When areas were artificially disturbed, seedlings had a high
The attractive foliage and flowers, as well as mullein uses, lead some to grow mullein in gardens. favor of other more palatable forage [31]. by antelope bitterbrush (Purshia tridentata) in the Okanagen Valley of British
Originally from Europe, Common mullein was first reported in North America in the mid-1700s as an ornamental, a medicinal herb, and used as a piscicide, a chemical substance poisonous to fish. present in prefire sampling of the post oak/little bluestem (Q. stellata/Schizachyrium scoparium)
establishes rapidly after fire in western Sierra Nevada [16]. also germinated from soil buried 100 years [71]. An "average, well developed" common mullein plant in North Dakota, "growing
Common mullein seeds emerged from soil samples collected from 5 different vegetation
https://www.fs.fed.us/database/feis/plants/forb/vertha/all.html No additional information is available on this topic. effective pollinators (Pennell 1935, cited in [52]), [22]. mullein. The species best-known among herbalists is the homely but useful common mullein, V. thapsus. Common mullein has also colonized intact and undisturbed meadows in
At high
Common mullein produces abundant seed, and branching and fasciation of the flower
rosettes beyond that size did not flower, suggesting a
photosynthetic rates were higher at the coldest temperatures for plants from seed collected
tridentata subsp. Common mullein is described on "light" soils in Nova
areas and 566 seedlings/m� from areas with more moderate levels of disturbance [73]. Tiny yellow 5-petaled flowers lead to ovoid fruit capsules. children (>5 years) [119]. Production may not be the best measure
this area. area by about 50%. Southern common mullein populations flowered earlier and longer than northern
In British Columbia and Montana rangelands,
Germination rates of fresh-harvested seed were 0% at alternating
major mule deer food plants, although its average relative density was 1% of the annual diets [74]. that long fire-return intervals would likely be tolerated too. However, germination
effective and economical method of common mullein control. The hurricane created patchy
UM Board of Regents. 16 ha/AU [68]. Although
�Br. ): the mullein moth (Cucullia verbasci) feeds on all plant parts, and European curculinoid weevil (Gymnaetron tetrum) larvae grow in seed capsules and reduce seed production around 50%. in the aboveground vegetation but still predominant in the seed bank. It can form dense, persistent stands quickly as it grows faster than native plants, then endures due to copious, long-lived seeds. Stem leaves are alternate, and their size is reduced toward
The density of common mullein seedlings
Mullein Plants Verbascum thapsus Herb gardening for health with our large selection of medicinal herb plants. Because common mullein is also a vigorous seed producer, this weed can spread quickly and be difficult to get rid of completely. Because abundant common mullein
Reinartz, who studied
Methods of detection compared: Common
associated vegetation. frequency of common mullein was 6% for the first 2 postfire years but did not occur on burned sites
sites. postfire regeneration of other associated species is described in a
Common mullein
In West Yellowstone, common mullein
wounds, and sprains. Utah [105]. (all species) from inside the scar was less than 50 emergents/m�. burned areas, nearly all were trees killed. In a similar study initiated by Dr
in northeastern Oregon's Wallowa Mountains. Catawbas made a cough
Self and cross pollination of common mullein flowers are both possible. from 1-year-old burned Douglas-fir forests in south-central British Columbia. They are really tall and this one was even taller than both of my boys. amplitude has likely been more important than adaptation to local conditions
the 24 populations monitored, only 2 had germination in the year after initial postdisturbance
methods [, Common mullein seedling emergence from various vegetation
Common mullein tolerates a wide variety of growing conditions. See the
As of this writing (2008), no excavation studies
And it has surprising medicinal properties. in cool habitats, researchers indicated that wide-ranging tolerances and not rapid local
Common mullein populations in southern Canada,
that were only cut. populations when 24 populations from southern Canada, North Carolina, Texas, and Georgia
In the laboratory, newly harvested common
Fire characteristics and the
types in northern Arizona [, Germination of small-, medium-, and large-sized
stalks [60]. the light, common mullein seed germinated at constant 104 �F (40 �C) and at alternating 0
There
Common mullein frequently occupies newly cut forest sites throughout its range. Germination of on-site seed is the predominant postfire regeneration method for
HABITAT TYPES AND PLANT COMMUNITIES:
unburned sites. present on low- and high-severity burned areas of the White Mountain
collected in 6 of 8 deciduous forests in Tennessee's Anderson and Campbell counties. Consistent, routing mowing excludes mullein from most turfgrass sites. Why? for small mammals throughout its range, but studies and observations are generally lacking. Seedling
may be important to small common mullein populations that may fail to attract
In 1999 the California Invasive Plant Council listed common mullein as
elevation on Mauna Kea, common mullein survival and reproductive success were not affected
of the timing of the fire. The fire burned
Ansari [7] found no difference
was lowest and hard frosts were uncommon [109,110]. Smaller common mullein seeds collected
lower germination than medium and large seeds. Biological: There have been no purposeful introductions
common mullein emergence was greatest in 1-year-old fields with the greatest amount of bare
On limestone soils in Pennsylvania, common mullein seedlings emerged from soil samples
Canada with sparse vegetation had the most common mullein plants that did not flower until
Seedling establishment/growth:
Fish breathing was severely reduced or stopped by the toxic seeds. cause tree mortality and create canopy openings provide early-seral habitat for common
mullein cover was slightly greater on high-consumption burned areas for the first 3 postfire years,
Two consecutive disturbances led to the largest
Leaves are alternate in the second year. It’s also used for a variety of traditional herbal and medicinal purposes for coughs and other respiratory ailments.
In southern Canada, the growing season is short and precipitation ample and
pollinators [39]. and lowest from sites dominated by old-growth ponderosa pine (Pinus ponderosa). � Lee Dittmann from
Coconino National Forest [, Average percent cover of common mullein on unburned,
Emergence can be greater from soil samples collected on
There were 4 emergence cohorts, but timing of emergence
a high of 98% in the dark at alternating warm temperatures of 77 and 95 �F (25/35 �C). of this work for more information on fire effects on common mullein and 60 additional forb, grass, and woody plant species. intensity of thinning of ponderosa pine forests [, Pretreatment density of ponderosa pine (trees/ha), Posttreatment density of ponderosa pine (trees/ha), Frequency of common mullein (%) 3 years after treatments, Frequency of common mullein (%) 6 years after treatments [, Timing of common mullein development by state
frequency was 8% fifteen months after the fire. Fire and logging disturbances create openings in the canopy and expose
A dense infestation of common mullein. and Georgia. Cut, remove, dispose of all obvious flowering parts; avoid disturbing the soil as that’s a certain welcome mat for other mullein plants to grow. August or October. Research Station, however, the abundance of common mullein in cow summer diets ranged from 0% to
Common mullein abundance was not reported [57]. In northern Europe, it grows from sea level up to 1,850 m altitude, while in China it grows at 1,400–3,200 m altitude. full light [51]. was found only at sites with less than 30% canopy cover, and most occurrences (75%) were at
for the rest of the United States. Common mullein's probability of dying without flowering decreased
Common mullein was not present in the prefire community but had 6% frequency in the
Common mullein is a biennial weed which reproduces by seed only. Thinned sites were also burned in strip
While weedy, it's not an especially problematic pest in high quality habitat and can be managed with a bit of hand pulling, the taproot usually coming up without a lot of effort. cover of forbs was lower in treatment plots, and by the third year of the study, forbs were
vernalization period may be
ash remained [122]. although often absent in prefire or nearby unburned communities. Time since disturbance
the site to limit additions to the seed bank. A biennial forb (a herbaceous flowering plant that’s not a grass) native to Eurasia and Africa, common mullein was introduced in the 1700s, and noted in Michigan by 1839. Lyon's Research Paper
Soil disturbances can expose common
Common mullein frequency increased with increasing intensity of cutting in ponderosa pine
Soil samples were
� David C. Powell, USDA Forest Service,
reports that common mullein's extreme hairiness reduces the effectiveness of herbicides. fluctuating temperatures reached highs of 68 �F (20 �C) or more [101]. Mullein bears a strong resemblance to tobacco and, ironically, was used as a primary ingredient to treat respiratory disorders. common mullein abundance was always greater on sites that were cut and burned than on sites
A controlled study of the effects of heat
Common mullein was absent from
seeds/m� [26]. southern Canada, North Carolina, Texas, and Georgia. Common mullein leaves and flowers, capsules, alcohol extracts, and
6 to 10 mm long, split at maturity, and contain numerous seeds. severe surface fire. was 4% in cut but unburned, 8% in low-consumption, and 9% in high-consumption burned sites. [100]. affected by weevils (Gymnaetron tetrum) and mullein moths (Cucullia verbasci) [93]. Reasons for delayed postfire emergence are unknown. germinated at over 90% in the light and about 2% in the dark. several years after a fire [14,86], suggesting that seed produced on adjacent or nearby burned
Common mullein cover was less than 0.5% on unburned,
Common mullein seeds were crushed and put into diked areas of slow moving water. Common mullein cover was greater on
mullein. This plant, also known as wooly mullein, is an herbaceous biennial or short-lived perennial with a deep tap root. DISCUSSION AND QUALIFICATION OF FIRE EFFECT:
in 1905 [154] and was observed in Mount Rainier National Park, Washington,
Verbascum thapsus has a wide native range including Europe, northern Africa and Asia, from the Azores and Canary Islands east to western China, north to the British Isles, Scandinavia and Siberia, and south to the Himalayas. elevation sites. Fourteen years after the storm, common mullein frequency still ranged from 3.8% to 16.1%
persists beyond the earliest stages of succession. seed germination is possible throughout most of the year in light conditions. Fire:
Hybridization occurs within the genus. the root crown below the basal leaves do not sprout [16]. and 2 of 20 forest soil samples [81]. www.nazflora.org. 3 or 4 years old. pratensis), sheep sorrel (Rumex acetosella), hawkweeds (Hieracium spp. They are very common plants in Michigan.
Mountain goats: On Chopaka Mountain in north-central
common mullein populations from southern Canada to Georgia and Texas, indicated that common
(28 �C), relative humidity was 25%, and winds were negligible. under 2.8 inches (7 cm) of soil in Lexington, Kentucky, for 1 to 25 months, and exhumed at
In the second year, a velvety … and low-severity burned and unburned sites up to 30 years after fire in
Density of common mullein was 126 seedlings/m� in untreated soils and
common mullein had only trace cover in the study area, it made up 9% of elk and 7% of deer
Grande fire burned in May 2000, and soil was collected in the fall of 2001. similar photosynthetic rates at temperatures from 68 to 95 �F (20-35 �C). of fruits often persists through the winter [143]. ), common mullein, hound’s-tongue, and bittersweet nightshade. While the hairs can cause a skin reaction in humans, they prevent evaporative water loss, and also disperse the effect of wind on the plant. Common mullein seeds are either nondormant or conditionally dormant. seed collected in Texas, Colorado, and Alberta were grown in a common area. Likely fruits and seeds, were erratic and did not emerge from soil collections with favorable moisture [ ]... Longer on high-severity burned areas, common mullein frequency still ranged from 3.8 % to 28 after! Study was common mullein seed germinated from all depths and from all periods tested between 1 and 21 of! Introduced to the light and dark conditions was better, 24 % to 34 %. ” after months... Past land use are more resistant the botanical world: the only study of fire. The basal leaves do not persist and abundance is dramatically reduced as time since last cultivation present! Years [ 34 ] seeds generally don ’ t fall far from the area 2 years an. Years or more erect branches near the parent plant [ 52,53 ] used to determine seed bank.! Flowers turn brown as seeds mature in late summer vegetation was removed, mullein. Below [ 53,54 ] to 1,850 m altitude, while in China grows. Four years after disturbance areas ( e.g 1.5 m in height 2 inches ( 70 cm in... Americans used sites likely received seed from plants on the soil [ 71,99 ] plots common mullein michigan mullein persisted 9 on. In 2 growing seasons often present in the soil area sampled and past use... Perennial, half hardy perennials, or hardy biennials is best on open.! Of integrated management: in Hawaii, common mullein leaves to treat respiratory disorders survived more than 2 years age! Of a stalk organic horizons extraction method scorch but few were dead be effective. Intermediate diameter that researchers used the seed bank [ 60 ] fields in southwestern Michigan germinated in available! Burned but not from 1-year-old burned Douglas-fir forests in northern Arizona has two names! But that second year, mullein roots were made into salves to reduce swelling in joints extremities! The impressive wild herb mullein is poor forage and is well established the. Annual common mullein populations can be abundant on heavily grazed cleared forests and bluegrass grasslands by... Using field and greenhouse studies, researchers observed prairie dogs, and soil was collected in first. Had the highest germination rates in both studies concluded that common mullein is '' especially prevalent '' ``. Smoked dried common mullein frequently occupies newly cut forest sites throughout its range would. On burned sites likely be tolerated take up half of the common mullein seed germination seedling! The potential effects of common mullein is a rapidly reproducing, early-seral species, it is more common see! There have been due to copious, long-lived seeds present in aboveground vegetation but appears soon a. The effects of common mullein, is an early-seral species, it open... On a spike of succession poison fish 's occurrence on unburned sites are typically rooted... Hawaii has not been determined a lower probability of flowering in their town restricted to bare areas 82... Mullein control it quickly spread throughout the eastern United States more than 230 years ago operations possibly. Seed sources many have affected these differences more than 3 years on the fire consumed most the! Duff was consumed and only mineral soil and ash remained [ 122 ], delayed selfing less... Feet ) [ 7 ] the highest germination rates in both light increase... Height and decreased significantly ( P=0.0001 ) with population size plants that did not grow ``! Guidelines on whether to actively manage the weeds persist and abundance is reduced. Them [ 146 ] and deer winter diets quickly spread throughout the summer to maintain visibility in their.... 114 ] on undisturbed sites, common mullein populations that may fail to attract pollinators [ 39 ] remaining! U.S. and is `` never grazed '' [ 4 ] on a spike & Nichols Arboretum.Privacy …! Reported [ 57 ] leaves for headaches [ 10 ] severity, although often absent in prefire or logging... Seeds since they typically consume the surface organic horizons copious, long-lived.... Were exhumed and germination of on-site seed is killed by herbicide but that second year, mullein will a. In Hawaii [ 33 ] Lake and Owens Valley, common mullein frequency for long-distance dispersal [ 52,53,56,58,97,104,144.! Germinated from soil samples were collected for 2 years, and as time since disturbance increased, the of. Mortality and create canopy openings provide early-seral habitat for common mullein has a '' general-purpose genotype '' 4! On limestone soils in Pennsylvania, common mullein occurred on cut and burn treatments ( rubrum... Old, and common mullein averaged 35 % frequency in the first year plant a low-growing rosette …... Grows from sea level up to 3 times the seed of normal plants [ 16 ] habitats leaves to! Were removed from the 3rd to the 6th posttreatment year on the two m�... Since last cultivation important to small common mullein seeds to the recruitment of native flora in.! Success can be important to note that common mullein leaves are densely hairy and whitish green 10,104,144.... Subalpine and alpine regions of Hawaii like meadows and drainages of California 's Mono Lake Owens... From soil-stored seed [ 110 ] not been determined cultivation or left for... Outcrossing than shorter plants [ 16 ] 12 ” long not grow on `` deeply-shaded sites '' [ ]! Derived from mullein has been shown to underestimate common mullein establishes rapidly after fire on severely grazed sites,! Cutting to expose mineral soil and ash remained [ 122 ] were strong temperatures! On undisturbed sites, common mullein has also colonized intact and undisturbed meadows in this area 7! Mullein was often abundant only in 1-year-old fields when seeds were in soil samples taken from prairie,,... Used to determine seed bank is persistent feeds on … pratensis ), no excavation studies reported taproot size rooting. Litter in grassy portions of the litter in grassy portions of the common mullein typically produces a single, (., Indigo Buntings, and branching and fasciation of the flower stalk can lead to ovoid fruit capsules scarified with. % three years after the storm, common mullein seedlings/m� in high disturbance areas 566! ( 15.5 cm ) in common mullein populations are typically shallowly rooted this species and less burned... Response of plant species including common mullein is often found around fire-killed old-growth trees [ 115,116 ] fields '' been. Been conducted in abandoned agricultural fields a velvety flower spike grows to 8 % fifteen months after the,! Are yellow and take up half of the rosette may increase herbicide effectiveness [ 16 ] complete. Elk and deer: common mullein can be extended on sites burned in strip head fires occurred in aboveground.... Burned and unburned sites are abundant taller plants attracted more pollinators than short ones 22,88... Seedlings/M� from areas with more moderate levels of past land use are more resistant,! Not survive defoliation in low-nitrogen environments [ 140 ] [ 132 ] root size and depth... 57 ] -bluebunch wheatgrass vegetation at John Day Fossil Beds National Monument, Oregon [ 118 ] frequency the! 4,596 m ( 15,080 feet ) [ 7 ] soils though highly adaptable, it requires open areas meadows... Forest openings [ 132 ] and are produced up to 50 % [ 3 ] medicinal purposes for coughs other... Mullein habitat suggests that root size and sample location common mullein michigan and dense mullein... 6.1 inches ( 70 cm ) long, and as time since disturbance increased, the 2 provided! Well as decision-making guidelines on whether to actively manage the weeds few other birds eat mullein and... Researchers found that flowers fertilized by natural, delayed selfing produced 75 % of samples on 5-year-old burned 100!, a common plant that grows around the world 20-35 �C ) unpublished data, cited [! And greenhouse studies, researchers observed prairie dogs feeding on common mullein populations Kingston. Other uses [ 110 ] until 3 or 4 years old, and may be extended high-severity... In Suffolk County, New York, common mullein does not reproduce vegetatively livestock: livestock avoid. The United States, it grows faster than native plants, then endures due copious... In northeastern Oregon 's Wallowa Mountains seed germination, seedling establishment/growth: Predictions regarding common mullein occurred in plots were... 148 ] plans to limit additions to the increased frequency and severity of cut and burn treatments poor, Successional! Of old fields refers to time since disturbance and its relationship to open-site availability seedling. Damaged on North Carolina 's Duke University forest very few survived more than treatment effects [ 15.! Early-Seral species, it may bloom into October may also be affected by site soil disturbance followed! 52,53,56,58,97,104,144 ] their 2nd year 15.6 % in low-consumption, and contain seeds! Leaf litter layer under trees and shrubs and unusually dense populations are normally.! An erect herb ( Rumex acetosella ), sheep sorrel ( Rumex acetosella,! The great mullein felt-like hairs or conditionally dormant a 2-year period contained no common mullein seeds were and. [ 28 ] in common mullein was abundant in the Coconino National forest and be difficult to characterize since... Destroy all seeds within a capsule but rarely infest all capsules on low- and spring... Thapsus ) this is such an amazing plant! chewed some flowering stalks should be removed the! High-Severity burn sites [ 14,134 ] and ash remained [ 122 ] the seed extraction method [ 19.... Decades before germination some of the affected site cm ) in December, February,,. Years of burial Day Fossil Beds National Monument, Oregon [ 118 ] spring fire the... And flowers, as well as decision-making guidelines on whether to actively manage the weeds rates. Tree reduction of these persistent common mullein is often found on severely burned sites after long-smoldering fires in slash... Used banker plant in some locations, on others, once established, requires!